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DLPFC projects directly to the SC.

It seems unclear what is the original source of SC inhibition in preparation of anti-saccades. Munoz and Everling cite the supplementary eye fields (SEF), dorsolateral prefrontal cortex (DLPFC) as possible sources, and the substantia nigra pars reticulata (SNpr).

The supplementary eye fields (SEF) and dorsolateral prefrontal cortex (DLPFC) both project directly to the SC.